城市化对北京自生植物物种组成和多样性的影响

doi:10.11707/j.1001-7488.LYKX20250501

Abstract

Abstract:

Objective: This study aims to explore the impacts of rapid urbanization, environmental factors, and cultivated plants on the species composition, spatial distribution patterns, and diversity of spontaneous plants in urban plant communities, providing crucial scientific basis for constructing low artificial maintenance, near-natural vegetation communities in urban green spaces. Method: The distance from the city center and the percentage of impervious surface area were selected as key indicators of urbanization intensity. The cross zone method was used to select 3285 spontaneous plants plots and 657 soil plots within the sixth ring road of Beijing for plant investigation and soil physicochemical property determination. Result: A total of 191 spontaneous plant species were recorded in this study, belonging to 172 genera and 61 families, including 28 trees, 16 shrubs and 147 herbs. The species composition and life form of spontaneous plants showed early regeneration and succession of the spontaneous plant communities. Meanwhile, 337 species of cultivated plants were recorded, belonging to 252 genera and 85 families. It was found that higher urbanization intensity increased spontaneous woody plant diversity, while decreased herbaceous plant diversity. In addition, spontaneous woody plant diversity was significantly positively correlated with the soil water content (P=0.043), total phosphorus (TP) (P<0.05), total nitrogen (TN) (P<0.05), and cultivated woody plants diversity (P<0.001), while not significantly correlated with green space factors. However, spontaneous herbaceous plant diversity was significantly negatively correlated with the soil TP (P<0.05), organic matter (OM) (P<0.01), the cultivated herbaceous plants diversity (P<0.01), and the intensity of disturbance in green space factors (P<0.01). Herbaceous species were able to quickly colonize and became pioneer plants under frequent human disturbance, and reduced the relying on the soil water and nutrition, such as Humulus scandens. Among the influencing factors of green space factors, soil factors and the diversity of cultivated plants, spontaneous species composition was mainly influenced by intensity of disturbance (contribution=32.1%). Conclusion: By altering the environmental factors, soil characteristics and the cultivated plants diversity in urban green spaces, the species composition and diversity of spontaneous plants can be improved in different types of urban green spaces and areas with urbanization intensity, which is conducive to increasing the resilience and sustainability of urban green space landscapes.

Key words: spontaneous plants, diversity, species composition, cultivated and spontaneous plants, urbanization

CLC Number:

S732
Q958.15

Dingjie Zhao,Tao Sun,Baoquan Jia,Shouhong Zhang,Hang Xu,Qimeng Yang,Mingqi Sun,Yawen Xue,Baohua Liu,Zhiqiang Zhang. Effects of Urbanization on Species Composition and Diversity of Spontaneous Plants in Beijing[J]. Scientia Silvae Sinicae, 2026, 62(4): 118-129.

Figures/Tables 8

Table 1

Table 2

Fig.1

Fig.2

Table 3

Multiple regression of spontaneous plant Shannon-Wiener diversity index and Pielou evenness index of spontaneous plants related to environmental variables"

变量 Variables		自生木本植物 Spontaneous woody plant				自生草本植物 Spontaneous herb plant
		Shannon-Wiener 多样性指数^a Shannon-Wiener diversity index		Pielou均匀度指数^b Pielou evenness index		Shannon-Wiener 多样性指数^c Shannon-Wiener diversity index		Pielou均匀度指数^d Pielou evenness index
		系数Coefficients	P	系数Coefficients	P	系数Coefficients	P	系数Coefficients	P
绿地因子 Greenspace factor	绿地类型Greenspace type	–0.076	0.354	–0.043	0.618	–0.026	0.734	–0.155	0.020^*
	绿化覆盖率 Greenspace area percentage	–0.060	0.463	–0.084	0.341	0.001	0.991	–0.061	0.368
	木本植物冠层郁闭度 Woody plant canopy density	0.008	0.914	0.060	0.458	0.048	0.518	0.069	0.284
	植物群落结构Community structure of planted plants	0.043	0.631	0.150	0.108	–0.101	0.199	0.072	0.285
	人工干扰强度Intensity of disturbance	–0.042	0.642	0.029	0.759	–0.271	0.001^^	–0.188	0.015^*
	绿地坡度和朝向 Slope andaspect of greenspace	–0.007	0.927	–0.074	0.369	–0.094	0.193	–0.012	0.849
土壤因子 Soil characteristics	土壤含水量 Soil water content	0.166	0.043^*	0.157	0.066	0.014	0.856	0.065	0.349
	土壤密度 Soil bulk density	–0.062	0.425	0.072	0.373	0.19	0.009^^	0.081	0.196
	全磷 Total phosphorus	0.170	0.036^*	0.134	0.113	–0.174	0.026^*	–0.080	0.229
	全氮 Total nitrogen	0.185	0.042^*	0.106	0.286	0.029	0.739	0.002	0.980
	有机质 Organic matter	–0.219	0.013^*	–0.176	0.060	0.059	0.483	–0.252	0.001^^
	有效磷 Available phosphorus	–0.006	0.938	0.037	0.666	–0.070	0.372	0.017	0.808
栽培植物多样性 Diversity index of cultivated plant	木本栽培植物香农多样性 Shannon-Wiener of cultivated woody plants	0.368	<0.001^^^*	—	—	—	—	—	—
	草本栽培植物香农多样性 Shannon-Wiener of cultivated herb plants	—	—	—	—	–0.212	0.003^^	—	—
	木本栽培植物均匀度 Pielou of cultivated woody plants	—	—	–0.047	0.591	—	—	—	—
	草本栽培植物均匀度 Pielou of cultivated herb plants	—	—	—	—	—	—	–0.497	<0.001^^^*

Table 3

Table 4

Fig.3

Table 5

References 0

	鲍士旦. 2000. 土壤农化分析. 3版. 北京: 中国农业出版社.
	Bao S D. 2000. Soil and agricultural chemistry analysis. 3rd ed. Beijing: China Agriculture Press. ［in Chinese］
	常　满, 李晓鹏, 张　彤, 等. 成都绕城绿道自生植物扩散特征及与栽培植物的共存. 生态学报, 2025, 45 (18): 9052- 9065.
	Chang M, Li X P, Zhang T, et al. Spread characteristics of urban spontaneous plants and their coexistence with the cultivars along Chengdu ring road greenway. Acta Ecologica Sinica, 2025, 45 (18): 9052- 9065.
	贺士元, 邢其华, 尹祖棠, 等. 1984. 北京植物志. 北京: 科学出版社.
	He S Y, Xing Q H, Yin Z T, et al. 1984. Flora of Beijing. Beijing: Science Press.［in Chinese］
	李晓鹏, 董　丽, 关军洪, 等. 2018, 北京城市公园环境下自生植物物种组成及多样性时空特征. 生态学报, 38(2): 581−594.
	Li X P, Dong L, Guan J H, et al. 2018. Temporal and spatial characteristics of spontaneous plant composition and diversity in a Beijing urban park. Acta Ecologica Sinica, 38(2): 581−594. ［in Chinese］
	刘欣仪, 陈春谛, 毕凌岚, 等. 城市老旧小区植物多样性及其价值研究: 基于居民支付意愿法. 生态学报, 2025, 45 (18): 9092- 9101. doi: 10.20103/j.stxb.202407161668
	Liu X Y, Chen C D, Bi L L, et al. A study on plant diversity and its value in urban old residential areas: based on the willingness to pay method. Acta Ecologica Sinica, 2025, 45 (18): 9092- 9101. doi: 10.20103/j.stxb.202407161668
	骆沁宇, 张梦园, 李晓璐, 等. 北京城市绿地不同生境自生植物多样性特征及其功能性状组成. 生态学报, 2024, 44 (11): 4744- 4757. doi: 10.20103/j.stxb.202311022387
	Luo Q Y, Zhang M Y, Li X L, et al. Diversity characteristics and functional trait composition of spontaneous plants in different habitats of urban green space, Beijing. Acta Ecologica Sinica, 2024, 44 (11): 4744- 4757. doi: 10.20103/j.stxb.202311022387
	马　杰, 贾宝全, 张　文, 等. 北京市六环内城市森林结构总体特征. 生态学杂志, 2019, 38 (8): 2318- 2325. doi: 10.13292/j.1000-4890.201908.035
	Ma J, Jia B Q, Zhang W, et al. The characteristics of urban forest structure within the sixth ring road of Beijing. Chinese Journal of Ecology, 2019, 38 (8): 2318- 2325. doi: 10.13292/j.1000-4890.201908.035
	彭　羽, 王玟涛, 卢奕瞳, 等. 城市化景观格局对本土植物多样性的多尺度影响: 以北京市顺义区为例. 应用生态学报, 2020, 31 (12): 4058- 4066.
	Peng Y, Wang M T, Lu Y T, et al. Multiscale influences of urbanized landscape metrics on the diversity of indigenous plant species: a case study in Shunyi District of Beijing, China. Chinese Journal of Applied Ecology, 2020, 31 (12): 4058- 4066.
	张梦园, 李　坤, 邢小艺, 等. 北京温榆河−北运河生态廊道自生植物多样性对城市化的响应. 生态学报, 2022, 42 (7): 2582- 2592.
	Zhang M Y, Li K, Xing X Y, et al. Responses of spontaneous plant diversity to urbanization in Wenyu River-North canal ecological corridor, Beijing. Acta Ecologica Sinica, 2022, 42 (7): 2582- 2592.
	中国科学院中国植物志委员会. 1978. 中国植物志. 北京: 科学出版社.
	Editorial Committee of Flora of China, Chinese Academy of Sciences. 1978. Flora of China. Beijing: Science Press.［in Chinese］
	周彦旭, 宗　桦, 杨珍茹, 等. 2025,. 自生植物研究进展与热点. 应用与环境生物学报, https://doi.org/10.19675/j.cnki.1006-687x.2025,.01004.
	Zhou Y X, Zong H, Yang Z R, et al. 2025,. Research progress and hotspots ofthe spontaneous plants at home and abroad. Chinese Journal of Appliedand Environmental Biology, https://doi.org/10.19675/j.cnki.1006-687x.2025,.01004. ［in Chinese］
	Cervelli E W, Lundholm J T, Du X. Spontaneous urban vegetation and habitat heterogeneity in Xi’an, China. Landscape and Urban Planning, 2013, 120, 25- 33. doi: 10.1016/j.landurbplan.2013.08.001
	Chang C R, Chen M C, Su M H. Natural versus human drivers of plant diversity in urban parks and the anthropogenic species-area hypotheses. Landscape and Urban Planning, 2021, 208, 104023. doi: 10.1016/j.landurbplan.2020.104023
	Chang M Y, Luo X Y, Zhang Y R, et al. Land-use diversity can better predict urban spontaneous plant richness than impervious surface coverage at finer spatial scales. Journal of Environmental Management, 2022, 323, 116205. doi: 10.1016/j.jenvman.2022.116205
	Chen X S, Wang W B, Liang H, et al. Dynamics of ruderal species diversity under the rapid urbanization over the past half century in Harbin, northeast China. Urban Ecosystems, 2014, 17 (2): 455- 472. doi: 10.1007/s11252-013-0338-8
	Cheng X L, Nizamani M M, Jim C Y, et al. Using SPOT data and FRAGSTAS to analyze the relationship between plant diversity and green space landscape patterns in the tropical coastal city of Zhanjiang, China. Remote Sensing, 2020, 12 (21): 3477. doi: 10.3390/rs12213477
	Cheng X L, Padullés Cubino J, Balfour K, et al. Drivers of spontaneous and cultivated species diversity in the tropical city of Zhanjiang, China. Urban Forestry & Urban Greening, 2022, 67, 127428. doi: 10.1016/j.ufug.2021.127428
	Chi Y, Shi H H, Wang X L, et al. Impact factors identification of spatial heterogeneity of herbaceous plant diversity on five southern islands of Miaodao Archipelago in north China. Chinese Journal of Oceanology and Limnology, 2016, 34 (5): 937- 951. doi: 10.1007/s00343-016-5111-4
	Chollet S, Brabant C, Tessier S, et al. From urban lawns to urban meadows: reduction of mowing frequency increases plant taxonomic, functional and phylogenetic diversity. Landscape and Urban Planning, 2018, 180, 121- 124. doi: 10.1016/j.landurbplan.2018.08.009
	Doroski D A, Duguid M C, Ashton M S. Forest patch size predicts seed bank composition in urban areas. Applied Vegetation Science, 2021, 24, e12534. doi: 10.1111/avsc.12534
	Fang Y T, Yoh M, Koba K, et al. Nitrogen deposition and forest nitrogen cycling along an urban–rural transect in southern China. Global Change Biology, 2011, 17 (2): 872- 885. doi: 10.1111/j.1365-2486.2010.02283.x
	Gao Z W, Song K, Pan Y J, et al. Drivers of spontaneous plant richness patterns in urban green space within a biodiversity hotspot. Urban Forestry & Urban Greening, 2021, 61, 127098. doi: 10.1016/j.ufug.2021.127098
	Ginocchio R, León-Lobos P, Arellano E C, et al. Soil physicochemical factors as environmental filters for spontaneous plant colonization of abandoned tailing dumps. Environmental Science and Pollution Research, 2017, 24 (15): 13484- 13496. doi: 10.1007/s11356-017-8894-8
	Guo P P, Su Y B, Wan W X, et al. Urban plant diversity in relation to land use types in built-up areas of Beijing. Chinese Geographical Science, 2018, 28 (1): 100- 110. doi: 10.1007/s11769-018-0934-x
	Guo Q F, Brown J H, Valone T J, et al. Constraints of seed size on plant distribution and abundance. Ecology, 2000, 81 (8): 2149- 2155. doi: 10.1890/0012-9658(2000)081[2149:COSSOP]2.0.CO;2
	Hu S W, Jin C, Huang L, et al. Characterizing composition profile and diversity patterns of spontaneous urban plants across China’s major cities. Journal of Environmental Management, 2022, 317, 115445. doi: 10.1016/j.jenvman.2022.115445
	Jha R K, Nölke N, Diwakara B N, et al. Differences in tree species diversity along the rural-urban gradient in Bengaluru, India. Urban Forestry & Urban Greening, 2019, 46, 126464. doi: 10.1016/j.ufug.2019.126464
	Li X P, Fan S X, Guan J H, et al. Diversity and influencing factors on spontaneous plant distribution in Beijing Olympic Forest Park. Landscape and Urban Planning, 2019, 181, 157- 168. doi: 10.1016/j.landurbplan.2018.09.018
	Li X T, Jia B Q, Li T, et al. Studying the spatial evolutionary behavior of urban forest patches from the perspective of pattern-process relationships. Urban Forestry & Urban Greening, 2023, 81, 127861. doi: 10.1016/j.ufug.2023.127861
	Li X T, Jia B Q, Zhang W, et al. Woody plant diversity spatial patterns and the effects of urbanization in Beijing, China. Urban Forestry & Urban Greening, 2020, 56, 126873. doi: 10.1016/j.ufug.2020.126873
	Mueller-Dombois D, Ellenberg H. 1974. Aims and methods of vegetation ecology. New York: John Wiley & Sons.
	Murakami K, Maenaka H, Morimoto Y. Factors influencing species diversity of ferns and fern allies in fragmented forest patches in the Kyoto city area. Landscape and Urban Planning, 2005, 70 (3/4): 221- 229. doi: 10.1016/j.landurbplan.2003.10.019
	Oldfield E E, Warren R J, Felson A J, et al. FORUM: Challenges and future directions in urban afforestation. Journal of Applied Ecology, 2013, 50 (5): 1169- 1177. doi: 10.1111/1365-2664.12124
	Pielou E C. The measurement of diversity in different types of biological collections. Journal of Theoretical Biology, 1967, 15 (1): 177. doi: 10.1016/0022-5193(67)90048-3
	Pregitzer C C, Ashton M S, Charlop-Powers S, et al. Defining and assessing urban forests to inform management and policy. Environmental Research Letters, 2019, 14 (8): 085002. doi: 10.1088/1748-9326/ab2552
	Qian S H, Qin D Y, Wu X, et al. Urban growth and topographical factors shape patterns of spontaneous plant community diversity in a mountainous city in southwest China. Urban Forestry & Urban Greening, 2020, 55, 126814. doi: 10.1016/j.ufug.2020.126814
	Shannon C E, Weaver W, Wiener N. The mathematical theory of communication. Physics Today, 1950, 3 (9): 31- 32. doi: 10.4135/9781412959384.n229
	Tredici P D.2010,. Spontaneous urban vegetation: reflections of change in a globalized world. Nature and Culture, 5: 299–315.
	Vakhlamova T, Rusterholz H P, Kamkin V, et al. Recreational use of urban and suburban forests affects plant diversity in a western Siberian city. Urban Forestry & Urban Greening, 2016, 17, 92- 103. doi: 10.1016/j.ufug.2016.03.009
	Vakhlamova T, Rusterholz H P, Kanibolotskaya Y, et al. Changes in plant diversity along an urban–rural gradient in an expanding city in Kazakhstan, western Siberia. Landscape and Urban Planning, 2014, 132, 111- 120. doi: 10.1016/j.landurbplan.2014.08.014
	Wang H F, Qureshi S, Qureshi B A, et al. A multivariate analysis integrating ecological, socioeconomic and physical characteristics to investigate urban forest cover and plant diversity in Beijing, China. Ecological Indicators, 2016, 60, 921- 929. doi: 10.1016/j.ecolind.2015.08.015
	Xiao L, Wang W J, He X Y, et al. Urban-rural and temporal differences of woody plants and bird species in Harbin City, northeastern China. Urban Forestry & Urban Greening, 2016, 20, 20- 31. doi: 10.1016/j.ufug.2016.07.013
	Yan Z G, Teng M J, He W, et al. Impervious surface area is a key predictor for urban plant diversity in a city undergone rapid urbanization. Science of the Total Environment, 2019, 650, 335- 342. doi: 10.1016/j.scitotenv.2018.09.025
	Yang F P, Ignatieva M, Wissman J, et al. Relationships between multi-scale factors, plant and pollinator diversity, and composition of park lawns and other herbaceous vegetation in a fast growing megacity of China. Landscape and Urban Planning, 2019, 185, 117- 126. doi: 10.1016/j.landurbplan.2019.02.003
	Yang J, Yan P B, He R X, et al. Exploring land-use legacy effects on taxonomic and functional diversity of woody plants in a rapidly urbanizing landscape. Landscape and Urban Planning, 2017, 162, 92- 103. doi: 10.1016/j.landurbplan.2017.02.003
	Zhang D, Zheng H F, He X Y, et al. Effects of forest type and urbanization on species composition and diversity of urban forest in Changchun, northeast China. Urban Ecosystems, 2016, 19 (1): 455- 473. doi: 10.1007/s11252-015-0473-5
	Zhao D J, Sun M Q, Xue Y W, et al. Spatial variations of plant species diversity in urban soil seed banks in Beijing, China: implications for plant regeneration and succession. Urban Forestry & Urban Greening, 2023a, 86, 128012. doi: 10.1016/j.ufug.2023.128012
	Zhao D J, Yang Q M, Sun M Q, et al. Urbanization and greenspace effect on plant biodiversity variations in Beijing, China. Urban Forestry & Urban Greening, 2023b, 89, 128119. doi: 10.1016/j.ufug.2023.128119

绿地因子指标 Greenspace factor indicators	分类或定义 Classification or definition
木本植物冠层郁闭度 Tree canopy density （%）	低木本冠层郁闭度 Low canopy density：0≤CD≤25
	较低木本冠层郁闭度 Relatively low canopy density：25<CD≤50
	中木本冠层郁闭度 Medium canopy density：50<CD≤75
	高木本冠层郁闭度 High canopy density：75<CD≤100

植物群落结构 Community structure of planted plants	乔木?灌木?草地结构 Arbor-shrub-herb structure
	乔木?草地结构 Arbor-herb structure
	灌木?草地结构 Shrub-herb structure
	仅草地植被群落结构 Single-layered herbaceous structure

人工干扰强度 Intensity of disturbance	轻度干扰：即绿地边缘有围栏或绿化隔离带，人类较难进入的绿地，无法在此类型绿地中游憩或休息 Low disturbance level: greenspaces enclosed by fencing or vegetated buffer strips， preventing public entry for recreation or rest
	中度干扰：即绿地周围仅有较矮的围栏或无围栏，游客不宜进入 Medium disturbance level: greenspaces with low or no perimeter fencing， where public access is discouraged or physically restricted
	高度干扰：即绿地周围无任何围栏，可直接进入绿地休息或游憩 High disturbance level: greenspaces with no perimeter barriers， allowing and accommodating direct public access for recreation and rest

绿地坡度和朝向 Slope and aspect of greenspace	绿地有坡度朝阴（即朝北）Greenspace with slope facing shade （north-facing slope）
	绿地无坡度 Greenspace nearly no slope
	绿地有坡度并朝阳（即朝南） Greenspace with slope and sunrise （south-facing slope）

植物区系、来源和生活型 Plant flora, origin and lifeform		自生植物 Spontaneous plant		栽培植物 Cultivated plant
植物区系、来源和生活型 Plant flora, origin and lifeform		数量 Number	百分比 Percentage (%)	数量 Number	百分比 Percentage (%)
科 Family		61	—	85	—
属 Genus		172	—	252	—
种 Species	本土植物 Native plant species	134	70.2	180	53.4
种 Species	外来植物 Exotic plant species	57	29.8	157	46.6
乔木 Arbor species	本土植物 Native plant species	21	11.0	52	15.4
乔木 Arbor species	外来植物 Exotic plant species	7	3.7	43	12.8
灌木 Shrub species	本土植物 Native plant species	8	4.2	31	9.2
灌木 Shrub species	外来植物 Exotic plant species	8	4.2	56	16.6
草本 Herb species	本土植物 Native plant species	105	54.9	97	28.8
草本 Herb species	外来植物 Exotic plant species	42	22.0	58	17.2
1年生草本Annual herb species		70	36.9	64	19.0
多年生草本 Perennial herb species		97	50.8	103	30.6

生活型 Lifeform	物种 Species	本土 Native	外来 Exotic	落叶木本植物/1年生草本 Deciduous vegetation/ annual herbs	常绿木本植物/多年生草本 Evergreen vegetation/ perennial herbs	相对多度 Relative abundance (%)	相对频度 Relative frequency (%)	相对盖度 Relative canopy coverage (%)	优势度/ 重要值 Dominant index/ important value(%)
木本植物 Woody plant	忍冬 Lonicera japonica		√		√	47.05	34.59	—	40.82
	栾 Koelreuteria paniculata	√		√		32.70	17.97		25.33
	枸杞 Lycium chinense	√		√		26.92	16.04		21.48
	榆 Ulmus pumila	√		√		18.33	19.24		18.78
	桑 Morus alba	√		√		9.78	17.97		13.87
	构 Broussonetia papyrifera	√		√		10.66	10.86		10.76
	荆条 Vitex negundo var. heterophylla	√		√		14.36	2.26		8.31
	臭椿 Ailanthus altissima	√		√		5.74	9.59		7.67
	白蜡 Fraxinus chinesis	√		√		3.29	4.33		3.81
	槐 Styphnolobium japonicum	√		√		2.87	4.68		3.78
草本植物 Herb plant	黎 Chenopodium album	√		√		12.95	9.22	7.08	9.75
	狗尾草 Setaria viridis	√		√		7.45	5.00	4.41	5.62
	假还阳参 Crepidiastrum lanceolatum	√			√	6.31	5.23	4.24	5.26
	紫花地丁 Viola philippica	√			√	2.31	5.76	3.81	3.96
	苦荬菜 Ixeris polycephala	√		√		4.29	4.48	2.84	3.87
	独行菜 Lepidium apetalum	√		√	√	4.03	3.18	2.40	3.20
	葎草 Humulus scandens	√			√	2.56	2.37	1.84	2.26
	早熟禾 Poa annua	√		√		3.07	1.77	1.88	2.24
	打碗花 Calystegia hederacea	√			√	1.81	2.53	1.82	2.05
	蒲公英 Taraxacum mongolicum	√			√	0.60	3.79	1.77	2.05

环境因子 Environmental factor	解释度 Explains (%)	贡献率 Contribution (%)	F	P
绿地类型 Greenspace type	0.5	3.5	1.3	0.19
绿化覆盖率 Greenspace area percentage	0.3	2	0.7	0.764
木本植物冠层郁闭度 Woody plant canopy density	0.5	3	1.1	0.334
植物群落结构 Community structure of planted plants	0.7	4.3	1.6	0.056
人工干扰强度 Intensity of disturbance	4.9	32.1	11.2	0.002^**
绿地坡度和朝向 Slope and aspect of greenspace	0.7	4.8	1.8	0.052
土壤含水量 Soil water content	0.8	5.2	1.9	0.022^*
土壤密度 Soil bulk density	0.4	2.8	1	0.42
全磷Total phosphorus	1.1	7.1	2.5	0.004^**
全氮Total nitrogen	1.1	6.9	2.5	0.004^**
有机质Organic matter	1.4	8.9	3.1	0.002^**
有效磷Available phosphorus	0.4	2.5	0.9	0.512
栽培木本植物多样性Shannon-Wiener of cultivated woody plants	1.1	7.1	2.5	0.006^**
栽培草本植物多样性Shannon-Wiener of cultivated herb plants	0.7	4.8	1.8	0.038^*
栽培木本植物均匀度Pielou of cultivated woody plants	0.3	1.9	0.7	0.802
栽培草本植物均匀度Pielou of cultivated herb plants	0.5	3	1.1	0.322

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Effects of Urbanization on Species Composition and Diversity of Spontaneous Plants in Beijing

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